Genshin Impact All Dendroculus Locations

The grind is real and has been for over three years. Honestly, at this stage, Genshin is an addiction for me. It’s been a long time, my fellow Travelers. Almost everything in the game demands ample time, whether it’s weapon progression, character ascension, or exploring new regions.

Sometimes, I even mindlessly grind for things I don’t want. What if a new character requires these materials in the future? Thoughts like these live rent-free in my head. 

These grinding sessions may be a needless quirk of my Genshin habits, but one I’m sure many can relate to is hunting Oculis in new regions.

With Anemoculi, Geoculi, and Electroculi collected, Dendroculi are the collectibles of choice at this moment for most Genshin players, seeing as the full collection is finally available.

Throughout my Genshin journey, it has been the most time-consuming chore, far outweighing my hunt for all of the Aranara.

I had to do world quests, clear ruins, and progress through puzzles and various challenges to finally max out Sumeru’s Statue of Seven. However, the experience was worth it, and through my pain, I will help you ease the burden of this task.

In this Genshin Impact Dendroculus guide, I will show you my routes and share the steps you must take to get all the Dendroculi in the game. Whatever you prefer, coffee or energy drinks, snacks or waffles, keep them handy because it’s time to snatch those cabbage-looking Oculis.

Let’s Go Dendro Hunting!

Before I start sharing the routes, map, and spots, I have a few important things to discuss:

• The routes, map, and places are in an order I felt was more comfortable following. You’re free to pursue your own path. I’ve prioritized the Oculis, which were above the surface, over those which were underground.
• I’ve mentioned world quests that will lead to those specific Dendroculi.
• I highly recommend following Genshin Chronicle’s Dendroculus video guide for a proper walkthrough. Yours truly has poured serious efforts into curating the video guide.
• Use characters that either help to reduce your running or gliding stamina or have skills that boost dexterity. Sayu, Yelan, Keqing, Xiao, and Lynette come to my mind, to name a few.
• I highly recommend completing these world quests before you begin your venture: Aranyaka, Girdle of Sands,  Apocalypse Lost, Golden Slumber, The Dirge of Bilqis, Khvarena of Good and Evil, and Afratu’s Dilemma.

Dendroculus Locations

Before we start going into individual routes, here is a list of the routes I have created for you:

1. Devantaka Mountain (21 Oculi)
2. Gandharva Village, Sumeru City, Alcazarzaray and Mawtiyima Forest ( 26 Oculi )
3. Vimara Village, Apam Woods, and Ruins of Dahri ( 32 Oculi )
4. Yasna Monument, Pardis Dhyai, and Vanarana ( 31 Oculi )
5. Realm of Farakhkert and Gavireh Lajavard ( 36 Oculi )
6. Near the Valley of Dahri ( 15 Oculi )
7. Land of Lower Setekh and Aaru Village ( 31 Oculi )
8. Hypostyle Desert ( 28 Oculi )
9. Desert of Hadramaveth ( 51 Oculi )
10. Entire Sumeru ( All routes combined – 271 Oculi )

Devantaka Mountain Route

Let’s first start with the nearest mountain range to grab some Oculi easily.

#1 Devantaka Mountain (21 Oculi)

The Dendroculus is above the ground near a tree. You can teleport to the nearest waypoint and make a run for it

#2 Under The Lake

You can do a plunge attack from the branch near the Dendroculus to grab it. There are two paths via which you can reach the Dendroculus. One is from the waypoint, and the other is from the path continuing underground.

#3 Near A Metallic Gear

On the ground, near a metallic gear. You can walk to it from the previous Dendroculus or teleport to the nearest waypoint.

#4 Near the Aranara

You will find it near an Aranara if you’ve not interacted with them yet. The Dendroculus is on the top of a large stone.

#5 Clear the Withering

You need to clear the Withering and then apply Dendro to the Rocklifting branch to make the Dendroculus appear. After that, you can collect it.

#6 Vimana Agama Quest

The Dendroculus can be collected during the Vimana Agama quest’s Royinjan chapter, where you have to play the Vintage Lyre to open a path.

#7 On the Top of a mountain

The Dendroculus is on the top of the mountain inside a broken hut. You can either climb or relocate with Four-Leaf sigils. Defeat the Eremites housed there and get free materials.

#8 On the Top of a mountain

You can fly from the previous Dendroculus’ location or the Statue of the Seven to get here. It’s floating above a metallic gear near the gigantic Ruin guard.

#9 Inside A Decayed Tree Trunk

You can fly from the Statue of the Seven to reach the location and grab the Dendroculus inside the destroyed tree trunk. 

#10 Near A Malfunctioned Ruin Guard

It won’t bite you, don’t be scared. The Dendroculus is in a net near a ruin guard by the river bank.

#11 Inside Gigantic Ruin Guard’s Finger

Your flying skills will be put to the test while grabbing this Dendroculus. You can either climb the Ruin guard’s arm or teleport via Four-Leaf sigils behind the arm to reach the hand. After that, jump and glide towards the Dendroculus.

#12 Below the Gigantic Ruin Guard’s Shoulder

You can relocate via Four-Leaf sigils to the spot. Two ruin machines will spawn as you come near the Dendroculus. It’s up to you to engage in the fight.

#13 Below the Gigantic Ruin Guard’s Shoulder

You can teleport to the highest waypoint on the other mountain and glide towards it. Another way is to climb all the way up and glide down to grab the Dendroculus.

#14 Near a Small Monument

You need to progress through the Aranyaka quest chain to obtain “The Rhythm of the Beastly Trial” to remove the barrier. Equip your Vintage Lyre and play the melody near the flower.

#15 Atop the Mountain

The Dendroculus is on the top of the mountain on a tree. There will be an Eremite sitting in its shade, so be prepared to engage in a fight.

#16 Whopperflowers for the Win

The Dendroculus is inside a barrier. You can reach the place from the previous Dendroculus’ location. Pick the plants one by one, defeat the Whopperflowers, and the barrier will vanish. A dialogue option appears on suspicious plants.

#17 On A Tree

Continue from the previous Dendroculus’ location to get here. You can easily jump on the tree and grab the Dendroculus sitting on it.

#18 On the Top of a Mound

It looks like an Aranara hut. You can continue from the previous Dendroculus’ location and grab the Dendroculus sitting on the top of a dome-shaped mound.

#19 Vintage Lyre: Here we go again

Continue from the previous Dendroculus’ location and go near the flower adjacent to a rock. You need to equip the Vintage Lyre to play music to remove the rock. Another way is to walk over the rock to get the Dendroculus.

#20 Dendrogranum Challenge

Keep going and reach the spot where you will encounter a Dendrogranum challenge. Make sure you have a bow character to shoot the Dendro gates ( Valaya ). After you successfully complete the challenge, the barrier around the Dendroculus will be removed.

#21 Inside the Mountain near the Gigantic Ruin Guard

You have to go inside the mountain to get the Dendroculus. Take the path northwest to the Gigantic Ruin guard, or do the Vimana Agama quest’s “Jazari’s chapter” to unlock the area.

Teleport inside the area or take the path to reach the Bouncy mushroom below the Dendroculus location. Apply Electro on it and jump to reach the Dendroculus.

Gandharva Village/ Sumeru City/ Alcazarzaray/ Mawtiyima Forest Route

Here’s the link to the route on the official map.

#22 In the Cave’s Entrance

You need to progress through the Agnihotra Sutra quest to get this Dendroculus. It is behind a net of branches that’ll unravel themself during the quest.

#23 Near the Statue of Seven

You can either transmit via a Four-Leaf sigil or glide to the spot

#24 On the Top of A Big Tree in the Village

Climb the tree with the help of Four-Leaf sigils. The Dendroculus is on the top of the tree.

#25 Inside a Treehouse

Go inside the hut and climb the stairs to reach the Dendroculus.

#26 Dendro Stone

Go to the spot displayed and attack the Rocklifting branch with Dendro. The branch will reveal the Dendroculus.

#27 Under the Bridge

You can either summon a Four-Leaf sigil with the help of a Clusterleaf of cultivation nearby or glide under the bridge.

#28 At the Bottom of the Waterfall

The Dendroculus is in front of the waterfall. You can easily glide your way down to grab it.

#29 At the top of the tallest tree in Sumeru City

Feel free to consume stamina-reduction food before climbing. The Dendroculus is on the second-highest branch, and you can also go to the highest one to get an achievement.

#30  Floating in the air

You can glide down from the previous Dendroculus’ spot or go to the location, apply Electro on the Bouncy mushroom, and get the Dendroculus.

#31  Under a Cliff

You can relocate via the Four-Leaf sigil and glide towards the Dendroculus. Another option is to slowly climb the cliff and touch the Dendroculus.

#32  On a pillar under the ground

You need to progress through the Agnihotra Sutra quest to obtain the Dendroculus. It can be obtained with or without the quest. Apply the Electro element to the Bouncy mushroom nearby and grab the Dendroculus.

#33 In a pit under the ground

You can continue from the previous Dendroculus to get here. Attack the Valaya, also known as Dendro gates with Dendrograna, and the barrier will vanish.

#34 In the Air

Go to the spot and apply the Electro element to the Bouncy mushroom.

#35 Behind the vines to an underground entrance

You need to progress through the Agnihotra sutra quest to remove the vines surrounding the entrance. Glide down to get the Dendroculus.

#36 Behind the vines to an underground entrance

Teleport to the nearest waypoint and glide towards the palace. You will end up on top of it.

#37 On top of a big mushroom

Teleport to the Statue of Seven and reach the location with the help of Four-Leaf sigils.

#38 On the top of a mushroom underground

Teleport to the waypoint in the vicinity of the Statue of Seven and make a run for the cave. Climb the tallest mushroom with the help of a Four-leaf sigil to reach the Dendroculus.

#39 On the road near the Statue of Seven

Teleport to the Statue of Seven and follow the road to the right. You will find the Dendroculus near an Abyss mage and a couple of Electro slimes. Weapons ready!

#40 Under a tree branch

Teleport to the waypoint near the Gandharva village and dash towards the small body of water. You will find the Dendroculus beside a Four-Leaf sigil.

#41 On the Top of a mountain

Continue from the previous Dendroculus’ location and start climbing the mountain. There is a Clusterleaf of cultivation that you can activate by applying the Dendro element to use a Four-Leaf sigil.

#42 On top of a perpendicular rock

Continue from the previous Dendroculus’ spot and use the Four-Leaf sigil to reach the place. An Aranara is also here prior to the task of interacting with them.

#43 On the top of a big mushroom

You can glide from the previous Dendroculus’ location. The Dendroculus is on a small mushroom-like flora. Apply the Dendro element on the Clusterleaf of cultivation to summon a Four-Leaf sigil to go to the next location.

#44 Dendrograna puzzle

Grab the Dendrograna and shoot towards Dendro gates ( Valaya ) to remove the barrier.

#45 Flying high in the air

Apply the electro element to the Tri-Yana seed at the mentioned location. Doing so will activate a wind current that will take you to the Dendroculus.

#46 Under a big mushroom

Teleport to the waypoint and glide under the mushroom. You can also use a Four-Leaf sigil to get to the Dendroculus.

#47 Under the ground, hidden by a Dendro mechanism

Teleport to the nearest waypoint and glide down to the cave. Once you reach the spot, you can activate the Rocklifting branch. For this, you have to progress through the Agnihotra Sutra quest.

Vimara Village/ Apam Woods/ Ruins of Dahri

Here’s the link to the route on the official map

#48 In A Cave

You can jump down from the road, make a 180-degree turn, and go inside a cave where a hilichurl is digging soil.

#49 On a cliff

It’s near the edge of a cliff by a Hilichurl. The hilichurl’s fate is in your hands, Traveler.

#50 Floating above a river

You can either summon a wave rider or freeze the water’s surface with a Cryo character like Kaeya. Alternatively, you can also use Kamisato Ayaka, Furina, or Mona’s alternate sprint. It’s up to you.

#51 In a Tree trunk

You can glide from the waypoint to a trunk near a shabby building.

#52 In a destroyed tree trunk

You can continue from the previous Dendroculus’ location or glide from the waypoint. There’s also a Four-leaf sigil that you can use to move.

#53 On top of a hut

The Dendroculus can be grabbed by climbing the hut or gliding from the waypoint

#54 Floating above a rusted gear

Go from the waypoint towards the location and fetch the Dendroculus.

#55 In the air

You need to apply the Electro element to a Bouncy mushroom to reach the Dendroculus in the air.

#56 Floating below a tree branch

Make use of the Four-Leaf sigil and glide down towards the Dendroculus

#57 Under a bridge

You can either glide down from the bridge ( needs solid flying skill ) or apply the Electro element to a Bouncy Mushroom below the bridge.

#58 Destroy the Withering

Equip your best team because the Withering would be a challenge after that, apply Dendro to the Rocklifting branch to reveal the Dendroculus.

#59 Near an entrance

The entrance is not under the tree. It’s near the roots and can easily be spotted.

#60 Below the Waterfall

Slowly glide down the waterfall to get the Dendroculus.

#61 Below the Waterfall

Cleanse the Withering to get rid of enemies and rejuvenate the surrounding area. Apply the Dendro element to the Rocklifting branch to reveal the Dendroculus.

#62 At the bottom of the tree

The location is tricky to get to. There is a Four-Leaf sigil that you can take to get under the roots and come to the hidden place under the tree.

#63 Floating above a feeble current of water

The Dendroculus is floating in the air. You can activate a Clusterleaf of Cultivation to summon a Four-Leaf sigil and grab it. Be my guest if you want the easy way of gliding to it

#64 Floating above in the air

Teleport at the domain’s location and apply Dendro to the Clusterleaf of Cultivation nearby. Keep moving towards the Dendroculus with the help of Four-Leaf sigils.

#65 On top of a tree

Teleport to the waypoint, and apply Electro on the Bouncy Mushroom to grab the Dendroculus.

#66 Floating in a cave’s entrance

You need to progress through the Varuna Gatha world quest to unlock the area. You can grab the Dendroculus during the quest.

#67 Floating above the water near the tree

I suggest gliding from the rocks nearby to get the Dendroculus. You can also glide down from the branch above the Dendroculus.

#68 Cleanse the Withering

Clear the area from the Withering and apply the Dendro element to the Rocklifting branch to unveil the Dendroculus.

#69 Below a staircase

There will be Eremites near the platform. Ignore them and grab the Dendroculus below it.

#70 Under a Rocklifting Branch

Clear the area from the Withering and apply the Dendro Element to the Rocklifting branch

#71 On the top of a tree under the ground

You need to progress through the Varuna Gatha world quest to open this area. Go into the cave, climb the tree, or jump on Bouncy mushrooms to get the Dendroculus.

#72 Rocklifting Branch

Just apply the Dendro element to the Rocklifting branch, and you can get the Dendroculus.

#73 On top of the branches

Go to the nearest waypoint, clear the enemies, and climb the tangled branches to get the Dendroculus.

#74 The Rhythm of the Gloomy Path

Go to the location and play the Rhythm with the Vintage Lyre to enter the portal. The Dendroculus will be on the top of a rocky outgrowth.

#75 Underground

You can get this during the Varuna Gatha world quest. To get the Dendroculus, you need to solve the puzzle and drain the water to make the way.

#76 On top of a tree

Simply climb the tree to get the Dendroculus. Beware of boars; they might charge at you.

#77 Withering Area

You need to cleanse the Withering from the area and apply the Dendro element to the Rocklifting branch to unveil the Dendroculus.

#78 The Rhythm of the Gloomy Path

Equip the Vintage Lyre to play the song obtained during the Agnihotra Sutra questline and get the Dendroculus.

#79 Inside the Cave

You can get this Dendroculus during the Dream Nursery world quest. The Dendroculus will be in front of you. Follow the path to get to the cave.

Yasna Monument / Pardis Dhyai / Vanarana

#80 On top of a rocky outgrowth

Teleport to the Statue of the Seven and go inside a cave to grab the Dendroculus with a Four-Leaf sigil.

#81 In a small hut

Out in the wild. You can go inside the small hut to fetch the Dendroculus.

#82 Floating above a waterfall

Jump on the Bouncy Mushroom to grab the Dendroculus

#83 Under a Rocklifting Branch

Cleanse the area from the Withering to rejuvenate the Rocklifting branch and apply the Dendro element on it to reveal the Dendroculus.

#84 The Cave’s Entrance

Follow the road, and it will lead to a cave’s entrance.

#85 Surrounded by Starshrooms

The Dendroculus is surrounded by the Fungi. Enter the area with caution to avoid being overwhelmed by Dendroshrooms.

#86 Inside an Aranara hut

Teleport to the Statue of the Seven and go inside a cave to reach an Aranara hut.

#87 Inside a tree trunk

Teleport to the Statue of the Seven and go to the spot with the help of Four-Leaf sigils.

#88 Inside the Gloomy Path portal

The Dendroculus is located right in the Gloomy Path portal. You can unlock the portal via the Rhythm of the Gloomy Path and explore the area inside.

#89 Bouncy Mushroom

You don’t need to fight enemies; just apply the Electro element on the Bouncy mushroom and jump in the air.

#90 On top of a dead tree

You can glide towards the pole from a higher elevation or climb it. The choice is yours.

#91 Under the ground

You can drain the water by activating the Dendro monument. The Withering affects the area, so cleanse it first to avoid a debuff. The Dendroculus is near a Claustroflora.

#92 Inside the area, near a small construct

Continue from the previous Dendroculus location. The Dendroculus is in the path.

#93 Inside a cave near hydro slimes

Teleport to the Statue of Seven and jump down to go inside the cave to get the Dendroculus.

#94 Bouncy Mushroom

Apply the electro element on the Bouncy mushroom to make your character jump high in the air.

#95 Another Bouncy Mushroom

Again, apply the Electro element on the Bouncy mushroom to grab the Dendroculus.

#96 Yet Another Bouncy Mushroom

Go below the bridge and apply the Electro element on the Bouncy Mushroom.

#97 Near a big mushroom under the ground

You need to complete the Phantasmal gate challenge to remove the barrier and glide towards a mushroom. The Dendroculus is floating above a tree trunk.

#98 Rocklifting Branch

Cleanse the Withering from the area and apply the Dendro element on the Rocklifting branch.

#99 Activate Dendro monuments

Go inside the mountain to activate four Dendro totems to unlock a mechanism to drain clogged water. Once you do that, unlock the area where the Dendroculus is.

#100 On a tree’s branch underground

Continue from the previous Dendroculus’ location, and you will find the Dendroculus floating on a branch.

#101 Bouncy Mushroom

Go to the spot and apply the Electro element on the Bouncy Mushroom to jump high in the air.

#102 Time to Jump

Take the help of the Four-Leaf sigil to reach the Bouncy mushroom, and apply the Electro element on it before you jump.

#103 Rocklifting Branch

Teleport to the Statue of the Seven and apply the Dendro element on the Rocklifting branch to reveal the Dendroculus.

#104 On top of a twisted tree

You can either climb the tree or glide from the nearest waypoint towards the spot. The choice is yours to make, Traveler.

#105 On top of a tilted monument

Teleport to the nearest waypoint and glide towards the Dendroculus’ location.

#106 Valaya Challenge

Equip the Vintage Lyre to play “the Rhythm of the Sprout” on the Dendrogranum branch and hit the Valayas with charged attacks of a Bow character.

#107 On a pile of stones

Follow from the previous Dendroculus’ location and climb the stones to get it.

#108 On a tree

Go to the spot and climb the tree to get the Dendroculus.

#109 Rocklifting Branch

You need to clear the Withering from the area and apply the Dendro element on the Rocklifting branch nearby to reveal the Dendroculus.

#110 On an Aranara tower

You can either glide from the nearest waypoint, climb the tower, or jump on a Bouncy mushroom to get to the top.

Realm of Farakhkert and Gavireh Lajavard Route

Here’s the link to the route on the official map.

#111 On top of a Hilichurl hut

Teleport to the nearest waypoint and climb the hut to get the Dendroculus. You can defeat nearby enemies to claim chests.

#112 On a cliff

I suggest teleporting to the domain to climb the cliff. The Dendroculus is floating on a cactus. Don’t worry; climbing the cactus won’t damage your character.

#113 On a wooden pole

Teleport to the waypoint near the cliff and climb the pole to claim the Dendroculus. You can unlock the chest by defeating the monster guarding it.

#114 In a cave

Teleport to the Statue of the Seven and make a run for the cave’s entrance. Glide down and use the Four-Leaf sigils to reach the Dendroculus.

#115 On a rock

The Dendroculus is located above the previous Dendroculus. Teleport to the Statue of the Seven and climb the protruding rock to grab it.

#116 In a canyon’s entrance

Teleport to the Statue of the Seven and go towards the canyon’s entrance. There’s a Four-Leaf sigil with which you can grab the Dendroculus.

#117 On a tree

You can easily reach the spot by initiating the challenge nearby to summon Four-Leaf sigils and make your way to the top. The Dendroculus is on a tree near a ferocious monster bird.

#118 On a tree’s root

The Dendroculus is on an overgrown tree root in front of the Statue of the Seven. Climb the root and claim it.

#119 Floating above in the air

You need to climb the mountain and interact with a special Four-Leaf sigil to claim the Dendroculus. Equip Sorush to place the special Four-Leaf sigil at an appropriate height and teleport to the Dendroculus.

#120 In a tree trunk

You can glide down from the previous Dendroculus’ location to the tree. The Dendroculus is inside the tree trunk’s opening. 

#121 Interact with the Environment

The Dendroculus is floating in the air, and you need a special Four-Leaf sigil to reach it. You can also glide towards it from a high elevation.

However, it’s easier to do with Sorush’s help. Equip Sorush, interact with Niroda fruit, and break orange rocks to reveal a special Four-Leaf sigil. You can reveal the platform and grab the Dendroculus with the sigil’s help.

#122 Amidst Four-Leaf sigils

Interact with a Clusterleaf of Cultivation near the water to summon a Four-Leaf sigil. You can reach the Dendroculus with this sigil’s help.

#123 Floating above an abandoned camp

You can glide towards the Dendroculus or use Gadgets like Red Feather Fan or Wind Catcher bottle to reach sufficient height and velocity.

Another way is to glide from the nearest waypoint towards the Dendroculus. I used Zhongli’s pillar, Venti’s wind current, and Kaedehara Kazuha’s skill from the waypoint and glided toward the Dendroculus from the waypoint on the bridge.

#124 Below a bridge

Teleport to the waypoint and glide down to grab the Dendroculus. It’s near a tree’s roots.

#125 On a tree

Continue from the previous waypoint and cross the bridge. You will see a tree where the Dendroculus will be floating on the tree.

#126 On a Treasure Hoarder’s Hut

Teleport to the waypoint near the big tree and go towards the cliff, where you will see the Dendroculus on a Treasure Hoarder’s hut. You can climb the hut to grab the Dendroculus.

#127 On a big tree with pink leaves

The Dendroculus is on a big tree. You can climb the tree to grab the Dendroculus.

#128 On a twisted branch

The Dendroculus is on a twisted branch. You can teleport to the waypoint on high elevation and glide towards it.

#129 On sharp tilted  branches

Teleport to the Statue of the Seven and use Sorush to place the special Four-Leaf sigil near the Dendroculus. Teleport and grab the Dendroculus.

#130 At the vortex’s entrance

The ambiance will change after you complete world quests. The Dendroculus is floating above an abyssal vortex. 

#131 On a cliff’s side

Climb the mountain and go near the devastated tilted branch to see the Dendroculus. Patiently glide towards it.

#132 Below a Cliff’s edge

Continue from the previous Dendroculus’ location to get this one. Glide down, and you will be able to get it. However, if you fall down or are at ground level, initiate the challenge nearby, and you will reach the top.

#133 Hovering over water near a riverbank

The Dendroculus is located at an isolated spot. It’s floating in the air near some trees and hilichurls. Easy to spot. 

#134 Near a Cliff

Equip Sorush and place the special Four-Leaf sigil at a better spot to grab the Dendroculus.

#135 On a Hilichurl hut

Teleport to the waypoint near the hut and reach the spot to clear the enemies and fetch the Dendroculus.

#136 In a Ruin construct

The Dendroculus is in a spherical ruin construct surrounded by Ruin machines. Be prepared to fight enemies when you reach the spot. 

#137 Below a Cliff

The Dendroculus is below a cliff. You can place the special Four-Leaf sigil with Sorush’s help. Once you do that, grab the Dendroculus.

#138 On a devastated/rejuvenated tilted branch

The Wooden branches will have a unique aesthetic depending on your world quests’ progress. The Dendroculus is out in the wild, sitting on the peak of a sharp wooden branch. 

#139 On an underground cliff

You need to progress through the Khvarena of Good and Evil world quest to unlock this area. Teleport to the waypoint under the ground and go towards the Dendroculus’ location.

#140 Under the Elevator

Heading down, you will encounter the Dendroculus below the Elevator. You can teleport, jump down, and glide towards the Dendroculus. You need meticulous gliding skills to get this one, so be a bit patient.

#141 Near a lava processing facility

The Dendroculus is floating near a lava processing unit under the ground. You can teleport to the underground waypoint and make a run for this one. You may need to climb the pipes surrounding the Dendroculus.

#142 Floating near metallic pipes

Teleport to the underground waypoint and head towards the spot. Utilize the wind current to get to the Dendroculus.

#143 Near Ruin guards

The Dendroculus is surrounded by Ruin guards. Progress through the Khvarena of Good and Evil world quest to get this one. 

#144 In an Arena

While following the Khvarena of Good and Evil world quest, you will find this Dendroculus in the final arena. Fend off the enemies, use the Four-Leaf sigils, and get the Dendroculus.

#145 Surrounded by roots under the surface

You need to teleport to the waypoint, break a large rock where a Hilichurl could be seen digging, set a Farrwick loose to make a path for you, and get the Dendroculus near a twisted root.

#146 On top of the spine

Depending on your world quests’ progress, the environment may be different. The Dendroculus can be grabbed via Four-Leaf sigils post-completion. Otherwise, the Dendroculus would be located on the top of a spine-like structure.

Valley of Dahri

Here’s the link to the route.

#147 Near a cliff

Climb the mountain and go towards the cliff where the Dendroculus is at.

#148 On a Fatui camp’s hut

You can teleport to the waypoint in the vicinity and glide towards the Dendroculus.

#149 Near the waypoint

Out in the wild. The Dendroculus is near the waypoint. There’s also a rock pile nearby that will give you 10K Mora

#150 On a small tree

Glide down from the waypoint, and you will find a tree surrounded by mushrooms. The Dendroculus is resting on the tree.

#151 On top of the Pyramid-like structure

You can dash to the top with characters like Yelan, Sayu, Lynette, or Xiao. The Dendroculus can be captured by gliding.

#152 Pyro totem puzzle

You need to apply Pyro on the totems to remove the invisible barrier around the Dendroculus. Equip a Pyro character and light the Pyro totems in any sequence.

#153 Inside a structure

Go to the entrance of the structure to initiate a mini-quest. You need to tail an Eremite to get the password. Once you do that, you will be able to open the gate and get the Dendroculus inside the structure.

#154 On a tree’s branch

Climb the mountain to go to the Dendroculus’s location. It is on the edge of a tree’s branch.

#155 On a massive rock

Teleport to the waypoint and make your way to the top with the help of Four-Leaf sigils. The Dendroculus is sitting on a tree’s branch.

#156 Near a big Ruin guard’s eye

You can glide down to the Ruin guard’s eye to obtain the Dendroculus.

#157 Inside the Ruin Golem

You need to initiate the world quest, Afratu’s dilemma. He will be near a small pond of water in front of the Ruin golem, where you can interact with him.

Once you initiate the quest, you will be able to unlock new waypoints and make your way to the Ruin Golem.

#158 Behind a Giant metallic gear

As you progress through the quest, Afratu’s Dilemma, you will hear a clanking sound. The Dendroculus is behind a massive metallic gear.

While you are in the cockpit, make sure to shoot all the shining locations displayed on the HUD.

Ruin Golem’s energy shots will land to open a path for the next Dendroculus.

#159 In a cave

You can get to the Dendroculus by utilizing the Four-Leaf sigil near it. I highly recommend doing the challenge nearby.

#160 Surrounded by the Withering

No need to cleanse the Withering. Go straight for the Dendroculus and jump forward to grab it.

#161 In a dungeon pit

There are multiple ways to reach the spot. However, teleporting to the Statue of the Seven and gliding down makes it easier to obtain the Dendroculus.

Land of Lower Setekh / Aaru Village

Here’s the link to the official route on the map.

#162 Near a small oasis

The Dendroculus is on a palm tree; you can glide towards it with the help of a Four-Leaf sigil.

#163 Inside a cave

You can enter the cave via a hole in the Dar Al Shifa area. Go inside the cave, break the rocks blocking the path, and you will find the Dendroculus hiding in a corner.

#164 Floating in a hole

The Dendroculus is right in front of the cave’s entrance, and you can jump inside the cave to get it.

#165 Near the Domain

Teleport to the domain, and you will see the Dendroculus on a small rock covered in sand.

#166 Near a chest

Take out your weapons, Traveler. The Dendroculus is near a chest guarded by Fungi.

#167 In the Aaru village

The Dendroculus is on a house in the Aaru village. Climb the house and jump to get the Dendroculus.

#168 Near a structure

The Dendroculus is near the waypoint in the Aaru village. Glide towards the Dendroculus to obtain it.

#169 In a hole in a cave

Continue from the previous Dendroculus’ location, and you will find the Dendroculus floating in a wind current. Make use of the Four-Leaf sigils and grab it.

#170 On a slanted rock

The Dendroculus is put in the wild. You can either glide towards it or climb the rock to fetch it.

#171 On top of the temple

You can either glide from the waypoint at a higher elevation or climb the temple. I suggest completing the Golden Slumber quest.

#172 Inside the temple

It’s best to collect the Dendroculus while you are doing the Golden Slumber quest, as post-completion, you will need maximum clearance on the Scarlet Sand slate to access the temple. The Dendroculus is located two levels deeper in the temple.

#173 Behind a statue

You need to go inside the building, pass through an invisible wall, and proceed through a challenge to get the Dendroculus. It may seem like a hassle, but it’s rather easy.

#174 On a vase in a corner

Go inside the building, unlock the doors, and you will see the Dendroculus in a corner behind a wall.

#175 In an archway

Inside the same structure, the Dendroculus is floating in an archway.

#176 On a wall

Teleport to the waypoint and climb the wall to grab the Dendroculus.

#177 On a tree stump

Teleport to the Statue of the Seven and glide to the tree stump to fetch the Dendroculus.

#178 Inside the structure

The Dendroculus is inside the structure. Press the red switch to access the area where it is.

#179 Beneath a cliff

The Dendroculus is located below a cliff. You can glide down to grab it.

#180 Up in the air

Take the wind current near the Dune to catch the Dendroculus floating above in the air. There will be a samachurl near the bottom the current of the wind.

#181 On a ledge

The Dendroculus is situated on a ledge under a cliff. Climb the cliff from the Statue of the Seven and glide down to grab it.

#182 Near a mountain

Teleport to the waypoint and go towards the Dendroculus’ location. It’s under the mountain behind three boxes. Break them, and you will be able to see the Dendroculus.

#183 Near the broken walls

The Dendroculus is outside the mausoleum, hiding behind broken walls surrounding the main building. 

#184 Above the temple

The Dendroculus is out in the wild. Teleport to the Statue of the Seven or continue from the previous Dendroculus’ location. You can easily run through it.

#185 On a headless statue

Where’s their head?!. The Dendroculus is sitting on top of a headless statue in the Khemenu temple. Teleport to the Statue of the Seven or continue from the previous spot to get it.

#186 On top of the building’s gate

Teleport to the waypoint and climb the building to get the Dendroculus. 

#187 Inside the building

You must progress through the Golden Slumber quest to get this Dendroculus. The Dendroculus can be obtained after using an elevator and completing a time trial challenge.

#188 On a cat-like statue

Proceed further into the building to get the Dendroculus sitting on top of a cat-like statue. Beware of enemies in the area.

#189 In the hallway

There might be enemies around you, so try to make a run for the Dendroculus to avoid a fight. The Dendroculus is on the floor.

#190 On a higher floor

You need to progress further through the Golden Slumber quest to access this level. The Dendroculus is on a lift’s platform.

#191 On a lift

The Dendroculus is on a lift’s platform that goes down after activation. It’s easy to spot.

#192 Floating in the lift’s path

You can get the Dendroculus just by riding the elevator. The Dendroculus is floating mid-air.

Hypostyle Desert

The link to the route on the map.

#193 Near a mural

Go inside the structure, turn right, and find the Dendroculus near a broken platform.

#194 Flying inside the structure

You need to activate the Anemo totems to summon a wind current. The Dendroculus is floating mid-air. Glide your way up to obtain it.

#195 Two levels below

The Dendroculus is two levels below the ground level. You need to drop down to obtain it.

#196 On a ship inside

The Dendroculus is on a ship framed inside the structure. You can get to it via the Four-Leaf sigils. You need to progress through the Golden Slumber quest series.

#197 Near a broken path

Go deeper into the underground area, and you will find a broken path. The Dendroculus is floating in the middle of it.

#198 In a wooden structure

Teleport underground and go to the spot where the Dendroculus is at. You can interact with the rock pile below to activate a wind current. You can also glide towards the Dendroculus.

#199 Enter the ruin

You need to clear the ruin to activate the last door to finally access the Dendroculus. There will be plenty of enemies, so equip your best team.

#200 Inside the structure

Enter the structure, interact with the mechanism to unlock the path, and jump down to get the Dendroculus.

#201 On a broken wall

Continue from the previous Dendroculus’ spot; this one is located deeper into the structure. You can also teleport inside the Dune of Magma to obtain the Dendroculus.

#202 Clear the ruin

Enter the ruin to solve puzzles to access an elevator. Once you finish the challenges, you will be able to ride an elevator that will take you to a room where the Dendroculus is located.

#203 On a mushroom head

The Dendroculus is deep inside the underground area. Glide towards the mushroom or make use of the Four-Leaf sigil to get the Dendroculus.

#204 Need full clearance

This one was a bit extreme in my opinion. You need full clearance on the Scarlet Sand slate to access the area with the tree. The Dendroculus is on a tree surrounded by the Withering.

#205 Floating in the air

Above the ground. The Dendroculus can be grabbed by activating an updraft.

#206 Solve a puzzle

It’s near the previous Dendroculus. It’s behind an invisible barrier, so you need to solve the puzzle nearby to get to it. 

#207 Behind a statue

In the underground area behind the statues. Make your way to the statues and use the Four-Leaf sigil to get to the top. The Dendroculus is hiding behind them.

#208 On a rock

This one’s easy. The Dendroculus is on a rock that you can easily climb

#209 On a statue below the surface

Teleport to the waypoint and go to the statue. There’s a Four-Leaf sigil that you can use to get to the Dendroculus.

#210 In the air

Teleport to the waypoint and use the updraft current to grab the Dendroculus.

#211 Inside the Pyramid-like structure

You need to complete challenges to unlock the elevator. Once you do that, you will be able to access the area with the Dendroculus.

#212 Inside the structure

Follow the Dirge of Bilqis world quest to access the area. The Dendroculus is to the right of the structure as you go inside.

#213 Interact with mechanisms

You need to interact with various mechanisms to unlock the area with the Dendroculus. Follow the direction of the Dirge of Bilqis’ quest and pay attention to tutorials.

#214 In a statue’s hands

Teleport to the waypoint and drop a few levels below to go to the statue. You can climb the statue to get the Dendroculus.

#215 On an underground pillar

Teleport to the waypoint under the surface. There’s a Four-Leaf sigil near the Dendroculus that can help you to get it. The Dendroculus is floating above the pillar.

#216 Blocked by an invisible wall

The Dendroculus is within your reach. Climb the sides to get to it. Don’t try to climb straight, as an invisible wall blocks it.

#217 Behind a construct

Teleport to the waypoint below the surface and go towards the big construct. The Dendroculus is behind the area.

#218 On a statue

Go inside the building and turn to the right, where you will see the Dendroculus near a statue’s head. Use the Four-Leaf sigil to attain sufficient height.

#219 On another statue

The Dendroculus is on another statue that has a Four-Leaf sigil above it. Make use of it and grab the Dendroculus.

#220 On top of the Mausoleum

Time to rub your hands and pack your water bottle. The Dendroculus is on the very top of the Mausoleum of King Deshret.

Desert of Hadramaveth

Take a look at the route here on the official map.

#221 Inside a building

You can open the door and enter the building. The Dendroculus is above a Four-Leaf sigil. Look up, and you will find the Dendroculus.

#222 Near small rocks

Teleport to the waypoint and glide towards the corner surrounded by rocks. The Dendroculus is out in the wild by a few bushes.

#223 On an upside-down pyramid-like platform

It’s in front of a large building hovering on a platform. You can glide to the Dendroculus to get it.

#224 On a broken wall

The Dendroculus is in plain sight. Teleport to the waypoint and rush towards the main building. Turn around, and you will find the Dendroculus.

#225 In a hidden cave’s entrance

You can get to the cave via the waypoint. It’s to the right of the waypoint. Make your way with the help of a Four-Leaf sigil.

#226 In the previous cave’s cavity

Roam in the cave where you collected the previous Dendroculus, and you will find a cavity with a Four-Leaf sigil. You can teleport via the sigil to grab the Dendroculus.

#227 On a small pillar

Glide down to get the Dendroculus sitting on a small pillar. There will be some enemies nearby, so beware of an ambush.

#228 On a statue

The Dendroculus is a bit lower in the depths of the valley. You can climb the statue and glide towards the Dendroculus to obtain it.

#229 Above an invisible platform

Equip a bow character and switch to aim mode to avoid falling off the platform. The area can be opened by following the Apocalypse lost world quest after you complete three other world quests: Tadhla the Falcon, The Falcon’s Hunt, and The Fallen Falcon.

#230 On a tree’s protruding roots

The Dendroculus can be grabbed by gliding towards it. It’s sitting on a tree’s roots.

#231 On a slanted rock

You can get the Dendroculus with the help of tall characters like Kaeya, Alhaitham, Zhongli, etc. Another way is to glide from the rock nearby to the Dendroculus.

#232 Inside a sandstorm

You must have a sufficient level on the Jinni in the magic bottle to dissipate the sandstorm. Progressing through the world quest series, Dirge of Bilqis will provide you with the gadget.

#233 On a tree’s branches

The Dendroculus is on a tree’s branches near rocks. Climb the rock, use the Four-Leaf sigil, and get the Dendroculus

#234 In the valley

Jump in the valley and glide down to get the Dendroculus. You can also glide from the ledge above it.

#235 Summon a wind current

Solve the Dendro totem puzzle to summon a wind current. Use Dendrogranum to break the Dendro rocks; a wind current will appear after you apply the Dendro element on Dendro totems. Use the updraft to fetch the Dendroculus.

#236 On a tree’s branch

You can glide towards the Dendroculus from a higher spot. The Dendroculus also has a Four-Leaf sigil near it.

#237 On a giant rock

You will come across the Dendroculus during the Dirge of Bilqis world quest. The Dendroculus is deep under the ground.

#238 Behind a Dendro rock

The Dendrogranum branch is near the Dendro rock. Use your elemental sight to scan the nearby area and attack the Dendro rock with the Dendrogranum to reveal the Dendroculus

#239 Floating in a pit’s entrance

It’s the best time to say hi to the Setekh Wenut monster. The Dendroculus is floating in the pit’s entrance. Teleport to the waypoint near the monster and get the Dendroculus.

#240 Behind a breakable rock underground

Teleport underground to find the breakable rock. Break the rock to unveil the Dendroculus.

#241 A level below under the ground

Teleport to the underground domain and jump a level below to turn right to find the hidden Dendroculus surrounded by the Fungi. The underground area is confusing, so take your time to explore.

#242 Near ruin guards

The Dendroculus is out in the wild. Go near the Ruin guard and jump to get the Dendroculus. You can also summon a Geo construct or wind current to grab it.

#243 Near a giant ruin robot

The Dendroculus is near an Eremite’s camp, chilling around the corner. Depending on your world quests’ progress, the Ruin robot might be present or not.

#244 On top of a ruin

Go to the spot, and you will find the Dendroculus sitting on a rock.

#245 In a Fatui hideout

The Dendroculus is outside the entrance to a Fatui hideout. You will visit the location again while doing the world quest: “Rejoice with me, for what was lost is now found”

#246 In a valley

You can get the Dendroculus by gliding from a higher spot.

#247 Near an underground waypoint

Apply Dendro on a Clusterleaf of cultivation behind the waypoint to summon a Four-Leaf sigil. Use the Four-Leaf sigils to go through the Dendroculus.

#248 On a small rock

The Dendroculus on a small rock near dried tree branches.

#249 Behind a Dendro rock

Grab the Dendrogranum from a branch nearby and attack the Dendro rock to reveal the Dendroculus.

#250 Near a shrine of depths

Climb the shrine and glide towards the Dendroculus to get it. 

#251 By a cliff

The Dendroculus is near a cliff’s side. You can glide down to obtain it. 

#252 Near a rock’s bottom

Again, glide down to get the Dendroculus. It’s in an open area near the bottom of a rock.

#253 Behind a Dendro rock

Use the elemental sight to scan for a Dendrogranum branch near the Dendro rock. Attack the Dendro rock with the Dendrogranum to reveal the Dendroculus.

#254 On a cactus

Teleport to the waypoint near the Dendroculus and climb the cactus to get the Dendroculus.

#255 Inside the Eternal Oasis’ area

Complete the world quest series, the Dirge of Bilqis, to unlock this area. The Dendroculus is inside a tree’s hollow trunk.

#256 Floating above a pond

Teleport to the Eternal Oasis’s area and get the Dendroculus floating above the pond’s water. You can walk on water here.

#257 Behind a tree

The last Dendroculus inside the Eternal Oasis. It’s behind a fallen tree, hiding in plain sight.

#258 In an Eremite hut

The area is roaming with Eremites. The Dendroculus is inside an Eremite hut. Enter the hut to increase your Dendroculus count.

#259 On a tree

The Dendroculus can be obtained by climbing the tree. 

#260 Use a wind current

The Dendroculus is floating high in the sky above an invisible platform. Use the elemental sight to find an Anemo totem nearby and activate a wind current that will lead you to the Dendroculus.

#261 On a cactus

The Dendroculus is on a cactus on the outskirts of the main desert. Climb the cactus to grab it.

#262 Above an invisible bridge

Go towards the Dendroculus. The Dendroculus is on an invisible platform that will help you to get it.

#263 Underground, Near the Giant ruin robot

Teleport inside the area and go towards the Dendroculus surrounded by the Fungi. It’s near a Primal Ember and is a part of the Dirge of Bilqis world quest. 

#264 In front of a mural

Teleport inside the area to go to the spot. The Dendroculus is a part of the world quest series Dirge of Bilqis. 

#265 In front of the giant robot

The Dendroculus is right in front of the giant robot’s eye. You can glide to the Dendroculus.

#266 On a tree

The Dendroculus is on a tree near a rocky stone. You can teleport to the nearest waypoint and make a dash towards it.

#267 On a rock

The Dendroculus is on a rock in front of the giant ruin robot. Climb the rock to get it. There’s also a viewpoint indicator at the same spot.

#268 Behind a Dendro rock

Break the spherical vortex around the Dendrogranum branch to get the Dendrogranum and attack the Dendro rock with it. Once you destroy the rock, the Dendroculus will spawn.

#269 In between some rocks

Teleport to the waypoint above the Dendroculus and glide down to get it.

#270 On an underground tree branch

Teleport inside the area and run towards the spot. You will find the Dendroculus sitting on a tree’s branch.

#271 Deep inside the underground area

The Dendroculus is on a rock under the surface. Teleport to the waypoint or continue from the previous Dendroculus’ location to get it. Make use of the Four-Leaf sigil to get to it.

Frequently Asked Questions

Congratulations, You Did It! 

So there you have it, guys, a complete and comprehensive Dendroculi guide, allowing you to max out Sumeru’s Statue of the Seven and get all the tasty rewards that come with that feat.

It’s been a grueling one for me, but now that I can take a step back and drink in every single Oculi discovery, I’m very proud, and I hope you will be too when you get to the end of this long and arduous task.

Now, if you don’t mind, I’m going to lie down and recharge. As always, thanks for reading Genshin Chronicle.